"Likewise, another study of 62-mile ultramarathon participants found that training volume and marathon personal record were the best predictors of ultra performance, far stronger than body size."
Of course they are! No one is suggesting that being bigger provides more of an advantage than training more. The question is whether, at a given training level, the same body types predict success in marathons versus ultramarathons. Logically speaking, you can't answer that question with a study that looks only at ultramarathoners (much less with studies of 17 and 19 participants, where a failure to find a correlation has a high risk of being a type II error).
As for the Millet et al. paper, it's from 2012, but he was presenting the same hypothesis at the Endurance Research Conference just last month. (And as I'm sure you know, Millet is probably the most prolific and respected ultra researcher out there over the past decade and a half. He's not just making stuff up.) As you point out, the leg muscle idea is nothing more than a hypothesis at this point, because the data to answer it one way or the other doesn't yet exist. But the studies cited in the RunnersConnect article certainly can't answer it -- especially since none of them even measured leg muscle mass.