Another thing is this: If you run the 400s with a true middle-distance stride, then there's more carryover to your performance at a mile.
If the 400s are a semi-sprint (and you're just recovering quickly), then you're not really developing the same specific skill of running that you'd use in a mile race. I suspect this was the situation with Clarke.
__________________________________________________________
On another note, if you have access to a treadmill I'd urge you to try uphill intervals. Not a terribly steep slope--maybe 5-10% (others have more experience with treadmills and might advise), because a really steep slope would, again, potentially mess with your mechanics. But the uphill work, in addition to developing your stride, seems to be kinder in terms of preventing injury (less impact).
Also, I know some people are substituting uphill walking for some or all of their long run. On a treadmill you can get a great aerobic workout just by walking (the slope would need to be kind of steep for this one--check your pulse to make sure it's at a productive level). I think one needs to build into this, though. You don't want to start with an hour going up a steep hill.