A couple of things I think are worth mentioning:
1. The increase in muscle tone from lifting can vary a lot.
The specific way you are designing and implementing a lifting session matters a ton.
On one extreme end:
-Reps to failure (even though the ideal distance to failure for strength is 3-5 reps)
-Differing exercises every 3-4 weeks,
-Lots of reps per set,
-Doing 1-2 really high volume lifting days,
-Really unstable exercises that are supposed to be more specific to running
-Periodizing lifting where you're only doing it half the year.
On the other end:
-5-7 reps from failure,
-Lower reps per set,
-Changing exercises when you stop improving (so likely after months rather than weeks)
-Aiming to get like 70% of the potential gains rather than 100%,
-Not being over concerned about lifts being hyper specific to running
-Spreading out lifting across more days if possible
Not going overkill with volume and realizing that quality is more important than quantity
-Lifting year round so your body can fully adjust.
If you're lifting is more like the second, the effect on muscle tone is going to be limited at most, whereas with the first you're going to feel terrible.
2. Sprints and strides can make some people feel better
An increase in muscle tone can be a good thing for some people. I personally will feel better after increasing my muscle tone from strides. I'm not that much of an outlier, this is pretty common in athletes whose best events are 800-3000. I feel best for a race by doing a bunch of really hard strides the day before and I'm really susceptible to over-tapering, which I think is related.
3. People generally overestimate how quickly they can adapt to something in the short term and underestimate the degree to which they can adapt to something in the long term
Well designed sprinting workouts designed to stimulate max speed are always low volume. It's common for people to see these workouts, want to see if they benefit from them, and then just copy them because it doesn't seem like that much work.
Like if someone saw that Cam Myers was doing 5 x 40m sprints twice a week, they might decide to start doing that workout, then when they copy that workout exactly, they get really sore, which is to be expected really. What if you had copied his threshold days, or even half of it, without any experience running.
If over an entire year you were to build to 200m of sprinting volume a week, starting at 10m, you'd be adding about 3.5 meters a week. It's unlikely that most people would so sore it ruins future Sub T sessions sprinting 10m once, then adding volume that slowly. I would bet a lot of money that people aren't doing that, which is strange, because most people know they can't jump straight into full double threshold with little running experience.