I was on a fairly successful college 4×1 team, and it was a wreck. We won virtually off talent alone. Three of the four runners on the relay were at best disinterested in the relay, or straight up didn't want to be there "because it didn't fit their training plan" (which is what our first leg was constantly complaining about).
One of them was a football player who hadn't run on a relay for a few years. He kept wanting to switch which hand he was holding the baton with DURING his leg "because he didn't like holding a baton in that hand; he ran slower with it in the hand he didn't like". When his teammates told him not to do that, he scoffed at us. We had to get the head coach to talk to him, and he was STILL was dubious, like we were the idiots. I think he still sneakily did it sometimes during his leg. He was our second leg.
The only one who was TRULY excited to run the relay was the slowest guy in the relay. Obviously he knew it was his ticket to do something special. However, even HE still wouldn't take instructions on the BEST way to hand off properly. He thought HE knew best...despite being the slowest guy of the group. He always tried to wait to hand the baton off to me until later in the zone, once I "got up to speed" (i.e. was about to run away from him). I was over half a second faster than him over 100 meters, but in his mind it was smarter to keep the baton in his hands a TINY bit longer (thus shorten my relay leg length) so I could "get up to speed". One time, he waited to long....and I ran away from him with no baton in hand. He was our third leg.
After some drama and a couple botched handoffs in meets, one of the coaches came up to me (me being the anchor and far and away the fastest guy of the team) and said, "listen, if you want to drop out of the relay just to focus on your own races as this is your last year, we'd understand. You've paid your dues. This relay could potentially win a title, but you're likely to win two individually anyway and we may not even be able to get the stick to you." I was tempted, but I figured it would be pretty dramatic in all the worst ways for the best guy on the team to simply leave the relay. So I said I'd prefer to stay on the relay.and just take our chances with the handoffs. No skin off my nose, really, anyway.
In the end, we won a title with that team, but of the nine races with that relay we ran that year, we DQ'd two of them, (once between leg two and leg three, and once from leg three to me) and had legitimately BOTCHED handoffs in another three. Those were the first five meets of the year. After that, we started to run some faster times, but the handoffs were still always a worry. In the last meet to win the title, I used to have it on video and again between leg two and three, they almost ran out of the zone again. Nobody ever talked about it, but I've seen it and it musta been CLOSE. Leg three went early and two had to REACH to get it to him.
The only thing that all made it work was the forced practice and failures, and even that didn't fully solve it with the poor egos, bad understanding of the concepts, and poor cohesion of the group overall. But we were so much faster than our peers, it didn't end up mattering. And it was STILL a question to the very end.
The real kicker? I think all those reps through the season put extra strain on my notoriously bad hammy. The relay with three guys who didn't even care about it won the title, and then I blew my hammy 30 meters into the 100 final and never ran fast again. Couldn't even get in the blocks for the 200 final, of course. While the guy who complained that the 4×1 practice didn't fit his training plan then got to bring up the rear of that 100 final and say he finally beat me in a 100. I still walked through the finish line in under 60 seconds, officially.
I hate that relay. But anyway, practice is the key. Even the idiots who don't play well together can pull it off with practice. Which will almost never happen with our top national guys, frankly.