It has always been obvious to me that genetics is the dominant influence on athletic ability. I spent my teenage years raging against my lack of running talent, utterly obsessed with the goal of becoming a world-class athlete when my genetics were average at best. Even back then I was clearly aware of the importance of genetics, but, being even stupider than I was slow, still held out for the laws of nature to make an exception for me, as though the reality of my genetic limitations would somehow magically disappear if only I could convince the running gods that I deserved it enough. In the end, I accomplished nothing in track, and became thoroughly depressed and maladjusted from having organized my life around unrealistic athletic endeavors at which I ultimately failed miserably.
Whoever denies the supremacy of genetics in running is clueless. It is the fundamental understanding upon which all other knowledge must be based. And if you do not clearly understand the role of genetics, you know literally nothing else about running, sport, or nature in general. It is THAT obvious.